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That is, you must be able to take a static photo of your hands and be able to tell what number is indicated, and be able via this process to count from 0 up to that number by 1s.
(The point of making the word "mean" bold is to emphasize it in speech: words are meant to be read, and we need to provide clues to readers as to how to read our words.)
The two prepositional phrases "in the number 1729" and "of the digits" are modifiers: "in the number 1729" tells us what digits we're talking about, and "of the digits" tells us what "each" refers to.
The core of the sentence is this:
You say "Each what?"; I say "Each digit." You say "Which digits?", and I say "digits in the number 1729."
In the end, however, you must answer the question
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What does #FFF8DC have to do with color?